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Rhinoceros Hornbill flying over misty Deramakot forest
Sabah, Malaysian Borneo
Deramakot
A working timber forest, and one of the world's great places to watch wild mammals.
Deramakot

A Forest,
Its Wildlife,
and the Travellers Who Found It

A short book on one of Sabah's most storied wildlife reserves.
Central Sabah  ·  Malaysian Borneo
Deramakot02
Foreword
Before You Begin
Aerial dusk view of the Deramakot base camp, Sabah Forestry DepartmentTom Cooper

This book moves in two parts. The first tells the story of a working timber forest that became, almost by accident, one of the world's finest places to watch wild mammals after dark. The second is a slower walk through the animals themselves — the mammals, birds and other wildlife that visitors most hope to meet on Deramakot's roads.

None of it replaces a guide. Deramakot is found by ear and by patient spotlighting as much as by knowing where to look, and the people who work its roads carry knowledge no book can hold. What follows is simply an introduction — enough to recognise what you're looking at, and why it matters that it's still there.

Foreword03
Contents
Inside This Book
Part One — The Forest
A Forest Ahead of Its Time05
From Timber to Trust06
Opening the Gates, 201407
The Night Jon Hall Came08
When Kate Humble Came Looking09
Built on Sleepless Nights10
A Circuit for Specialists11
Part Two — The Wildlife
The Mammals, Birds & Others13–35
Part Three — Practical
Planning Your Visit36
Field Checklist, 620 species37–43
References & Acknowledgements44
Contents04
Part One · Chapter One
A Forest Ahead of Its Time
Aerial view of Deramakot's forest canopy with mist over the hillsTom Cooper

In 1989, long before "sustainable forestry" was a phrase tourists cared about, the Sabah Forestry Department made a decision that would, decades later, reshape wildlife tourism in Borneo. Working with technical support from the German Agency for Technical Cooperation, the department began developing a management system for Deramakot Forest Reserve built around a simple but, at the time, unusual premise: that a working timber forest could be logged commercially and still function as a healthy ecosystem.

The approach became known as Reduced Impact Logging, or RIL — a method of harvesting timber designed to keep damage to the surrounding forest to a minimum. Only carefully selected trees above a minimum diameter are felled. Fruiting trees, a critical food source for the reserve's mammals and birds, are left standing. Buffer zones are maintained along rivers, watersheds and steep slopes.

Deramakot became the Sabah Forestry Department's flagship — a working demonstration of what steady political support and disciplined management can achieve.

This was not an easy or obvious choice for a state forestry department to make in 1989. It meant slower harvests and a longer road to profitability than conventional logging offered. But it set Deramakot on a trajectory that, a generation later, would make it famous for reasons nobody at the time anticipated.

Part One — The Forest05
Part One · Chapter One, Continued
From Timber to Trust

In 1997, Deramakot became the first natural forest in Southeast Asia to be certified as a "well-managed forest" by the Forest Stewardship Council. No tropical rainforest has held the certification for longer. It lapsed in late 2019 and was regained in 2024.

What followed was not something the department had explicitly set out to achieve, but it followed logically from the method. Researchers working in Deramakot began documenting healthy populations of animals that are usually the first to vanish from logged landscapes — Clouded Leopards, Sun Bears, Orangutans, and a striking diversity of civets — persisting inside an active, licensed timber concession.

Across much of Southeast Asia, "logged forest" and "wildlife-rich forest" were treated as opposites. Deramakot offered quiet evidence that with sufficiently careful management, they did not have to be.

Part One — The Forest06
Part One · Chapter Two
Opening the Gates, 2014
Aerial view of the winding gravel road through Deramakot Forest ReserveTom Cooper

By around 2014, the Sabah Forestry Department began permitting a small number of licensed operators to bring visitors into Deramakot, using the department's own staff accommodation at the base camp as a starting point. It was a modest, cautious opening — nothing like the purpose-built ecotourism infrastructure found in Sabah's national parks.

There were no manicured trails, no visitor centre. Guests stayed in the same staff housing used by forestry personnel and joined slow, deliberate night drives along the reserve's forestry roads — a network of wide, well-kept gravel tracks left behind by decades of careful timber work.

That simplicity turned out to be an asset rather than a limitation. The roads were quiet, wide enough for good sightlines, and ran straight through forest the animals actually use.

Part One — The Forest07
Part One · Chapter Two, Continued
The Night Jon Hall Came

By late 2014, reports of Deramakot had begun circulating within a small, tightly networked international community: mammalwatchers, whose travel is built specifically around finding wild mammals. Early accounts described Sun Bears, Banded Palm Civets and Marbled Cats turning up within days.

In May 2015, Jon Hall arrived — founder of mammalwatching.com and widely regarded as the community's most prolific chronicler. He travelled with fellow enthusiasts Tomer Ben-Yehuda and Jean-Michel Bompar, guided through the operator Adventure Alternative Borneo.

Over four nights in May 2015, the group recorded thirty-one mammal species — Binturong and Banded Palm Civet among them.

In a community that relies heavily on shared trip reports to decide where to travel next, that account functioned as an introduction. Deramakot's status as one of the world's premier mammal-watching destinations traces directly back to that period.

Mammalwatching.com has since become the place where Deramakot trip reports are shared, and each new report has added to the reserve's reputation among the people most likely to make the journey.

Jon Hall, founder of mammalwatching.com, speaking at a conference1Stop Borneo
Part One — The Forest08
Part One · Chapter Three
When Kate Humble Came Looking

The BBC presenter and naturalist Kate Humble arrived at Deramakot in January 2019 with a more pointed question than most visitors bring: could a forest that still earned its keep as a working timber concession really hold onto the wildlife everyone assumed logging destroyed, or was that just a good story? She spent five days driving the reserve's forest roads to find out.

The animals obliged steadily rather than spectacularly — Slow Lorises, flying squirrels, Leopard Cats, Sun Bears and Bornean Pygmy Elephants, with palm civets and Banded Civets turning up so often she came to think of them as regulars. An Otter Civet, rarer by her own reckoning than the cat everyone comes for, was the trip's real prize.

The Sunda Clouded Leopard eluded her, as it eludes many visitors.

The Clouded Leopard itself never appeared. Humble left anyway believing she had her answer: that a forest can pay its way in timber without being stripped of the wildlife that makes it worth protecting in the first place — a conclusion delivered, fittingly, to the sound of gibbons singing at dawn.

Part One — The Forest09
Part One · Chapter Four
Built on Sleepless Nights

In the years since 2015, Deramakot's reputation has continued to build. Species once treated as a once-in-a-lifetime encounter anywhere in Borneo — Marbled Cat, Sun Bear, Otter Civet, Binturong, the Sunda Clouded Leopard — are today reported with a frequency that would have seemed implausible before the reserve opened to visitors.

The format that made this possible — slow night drives, patient spotlighting, extended multi-night stays rather than single-day visits — has since been adopted, in varying forms, by other Sabah forest reserves seeking to open their own land to wildlife tourism.

A forestry department's 1989 commitment to sustainable timber management has grown, three decades on, into one of the clearest examples in the region of conservation and tourism reinforcing one another.

Part One — The Forest10
Part One · Chapter Four, Continued
A Circuit for Specialists

What began with a handful of independent mammalwatchers has, in the years since, become a fixture on the calendars of specialist wildlife travel companies around the world. Some now build whole itineraries around a week at Deramakot, timed to the reserve's spotlighting rhythm rather than a conventional daytime safari.

Small-group departures now run every year, aimed squarely at the reserve's specialist rarities, and the trip reports they leave behind — some logging more than forty-five mammal species in a single visit — have become one of the best running records of what the reserve actually holds.

None of this existed before 2015. That a working timber concession can now draw visitors back year after year is itself a kind of finding — proof, gathered one trip report at a time, that the approach Deramakot adopted in 1989 works.

Part One — The Forest11
A forestry road through Deramakot Forest Reserve — the kind of track most night drives follow.
Banded Civet Hemigalus derbyanus
Endangered  ·  Nocturnal  ·  ground-dwelling

Unmistakable among Deramakot's civets, the Banded Civet carries bold dark bands across a pale cream coat, a pattern that breaks up its outline in the dappled beam of a spotlight. It roots earthworms and invertebrates from the leaf litter with an elongated, mobile snout.

Sightings typically come low to the ground, often crossing the main road or foraging just inside the treeline — moving alone, with a distinctive low, searching gait.

SizeHouse-cat length, slimmer
DietEarthworms, insects, small animals
ActiveNight
Where to lookOn the ground: road edges and leaf litter
Part Two — The Wildlife13
Malay Civet Viverra tangalunga
Least Concern  ·  Nocturnal  ·  common

The largest and most frequently encountered of Deramakot's terrestrial civets — a heavily built, grey-brown animal marked with dark spots and bands and a strongly ringed tail. A generalist, taking fruit, small vertebrates and invertebrates as available.

Comfortable in disturbed and regenerating forest, it is one of the more dependable sightings on a night drive, usually crossing open road sections at a steady, confident pace.

SizeLarger than a house cat
DietFruit, insects, small animals
ActiveNight
Where to lookCrossing the road and along forest edges
Part Two — The Wildlife14
Bornean Striped Palm Civet Arctogalidia stigmatica
Least Concern  ·  Nocturnal, arboreal  ·  occasional

A slender, fully arboreal civet named for the three faint dark stripes running down its back. Unlike the ground-foraging civets on the previous pages, it rarely descends to the road, instead moving through the mid and upper canopy along tangles of liana and branch.

Sightings are usually a pair of eyeshine high in the canopy, followed by patient tracking to confirm the animal's outline and tail. Largely frugivorous, and often located near fruiting fig trees.

SizeHouse-cat size, very long tail
DietMostly fruit, especially figs
ActiveNight
Where to lookHigh in fruiting trees
Part Two — The Wildlife15
Part Two · The Wildlife
Two More Civets

Otter Civet

Cynogale bennettii  ·  Endangered

One of the rarest and strangest civets in the world, with webbed feet, a broad flat muzzle and dense whiskers, built for hunting fish, crabs and frogs along streams. It is so seldom seen anywhere that a sighting is the highlight of a trip, and Deramakot is one of the very few places where visitors have a realistic chance.

SizeHouse-cat size, short tail
DietFish, crabs, frogs
ActiveNight
Where to lookStreams and wet gullies by the road

Island Palm Civet

Paradoxurus philippinensis

Often the most frequently seen civet on a night drive: grey-brown with darker legs, faint spots and a pale face mask. It spends much of its time in trees eating fruit, and is regularly found in fruiting trees beside the road, sometimes several together.

SizeHouse-cat size, long tail
DietFruit, insects
ActiveNight
Where to lookFruiting trees beside the road
Part Two — The Wildlife16
Leopard Cat Prionailurus javanensis
Least Concern  ·  Nocturnal  ·  common

Roughly the size of a domestic cat, with a spotted golden coat, the Leopard Cat is Deramakot's most widespread small felid and often the first cat species a visitor records. It hunts rodents along road edges and regenerating forest patches, habitat that reduced-impact logging leaves in good supply.

Its eyeshine is bright and low to the ground — usually two points of light before the animal's full outline resolves in the beam.

SizeDomestic-cat size
DietRats, frogs, small birds
ActiveNight
Where to lookRoad verges, low in the beam
Part Two — The Wildlife17
Sunda Clouded Leopard Neofelis diardi
Vulnerable  ·  Mostly nocturnal  ·  rare, increasingly reported

The largest cat in Borneo, and the animal most visitors hope for above all others. Named for the large, irregular cloud-shaped markings across its coat, it is equally at home on the ground and in the canopy, with proportionally the longest canine teeth of any living cat. Despite the name it is not a leopard at all, but a separate, ancient branch of the big-cat family.

Camera-trap surveys run between 2007 and 2023 across Deramakot and its neighbouring reserves identified 52 individual adults and recorded the oldest wild Sunda Clouded Leopard ever documented — a female estimated at roughly eight and a half years old. The same research found females about 68% harder to detect than males, likely because they spend more time away from the roads and trails where cameras are set.

Once considered close to mythical in Bornean forests, it is now recorded here with genuine, if still infrequent, regularity — a shift widely credited to the reserve's well-managed forest and healthy prey base. Fewer than 10,000 mature individuals are thought to remain across Borneo and Sumatra combined.

SizeLarge-dog length, tail as long again
DietPrimates, deer, pigs and other mammals
ActiveMostly night; occasionally seen by day
Where to lookOn the roads, or in the trees above them
Part Two — The Wildlife18
Marbled Cat Pardofelis marmorata
Near Threatened  ·  Day and night, arboreal  ·  uncommon, well recorded

A small, exceptionally beautiful cat with a marbled coat pattern reminiscent of a miniature clouded leopard, and a tail often as long as its body. Highly arboreal, it uses that tail for balance while moving through branches after birds, squirrels and small mammals.

Few sites anywhere in its range offer the sighting frequency now associated with Deramakot, where it has become one of the reserve's calling-card species.

SizeHouse-cat size, tail as long as its body
DietBirds, squirrels, rats
ActiveDay and night
Where to lookMid-canopy branches near the road
Part Two — The Wildlife19
Yellow-throated Marten Martes flavigula
Least Concern  ·  Mostly diurnal, arboreal  ·  rarely seen

A long-bodied, glossy member of the weasel family, dark brown with a bright yellow throat and bib and a long black tail. It is as quick on the ground as in the trees, and hunts squirrels, birds and rodents, as well as taking fruit and honey. It is often found in pairs or small family groups.

Despite living across the reserve, the marten is rarely seen at Deramakot. It moves fast and keeps to cover, so most sightings are a few seconds long, a face looking down from a branch or a tail disappearing into the foliage. Treat one as a bonus, never as something to plan on.

SizeBigger than a house cat, long tail
DietSquirrels, birds, fruit, honey
ActiveDay
Where to lookCrossing the road, or in the canopy
Part Two — The Wildlife20
Part Two · The Wildlife
Bear and Bearcat

Sun Bear

Helarctos malayanus  ·  Vulnerable

The smallest of the world's bears, black with a pale muzzle and a cream or orange chest patch that is different on every animal. A strong climber, it tears into trees for honey and termites with long claws and a very long tongue. At Deramakot it turns up on night drives, sometimes high in a tree, sometimes crossing the road.

SizeSmallest bear: 25–65 kg
DietTermites, honey, fruit
ActiveMostly night here
Where to lookTree trunks, or crossing the road

Binturong

Arctictis binturong  ·  Vulnerable

Often called the bearcat, though it is neither: a large, shaggy civet with tufted ears and a thick tail it can grip branches with. It feeds mostly on fruit, figs especially, and moves slowly through the canopy. Spotlighting a fruiting fig at night is the best way to find one, and it often stays put long enough for a good look.

SizeBody up to about 90 cm, tail almost as long
DietMostly figs and other fruit
ActiveMostly night
Where to lookFruiting figs in the canopy
Part Two — The Wildlife21
Mouse-Deer Tragulus napu & T. kanchil
Least Concern (both)  ·  Nocturnal / crepuscular  ·  very common

Deramakot hosts both the Greater and Lesser Mouse-deer, among the smallest hoofed mammals on Earth, distinguished mainly by size and throat markings — delicate, deer-like animals standing barely knee-high, with slender legs and small tusk-like canines rather than antlers.

Frequently the first mammal of the night, often frozen at the road edge in the beam before bolting for cover — a reliable warm-up before the reserve's rarer specialists appear.

SizeRabbit to hare size
DietFallen fruit, leaves, shoots
ActiveNight and dusk
Where to lookRoad edges, frozen in the beam
Part Two — The Wildlife22
Part Two · The Wildlife
On the Forest Floor

Bornean Bearded Pig

Sus barbatus  ·  Vulnerable

A tall, lean wild pig with a long head and a bushy pale beard along the snout. It roams widely for fallen fruit, seeds and roots, and in big fruiting years groups can travel long distances following the food. In 2021 African swine fever reached Sabah and killed very large numbers of Bearded Pigs across the state. They are now coming back at Deramakot, and family parties are again being seen along the roads.

SizeBig males over 100 kg
DietFallen fruit, seeds, roots
ActiveDay and night
Where to lookAlong the roads and forest edges

Malayan Porcupine

Hystrix brachyura  ·  Least Concern

A large, ground-dwelling rodent armed with long black-and-white quills, which it raises and rattles when threatened. It forages at night for roots, fallen fruit and bark, and spends the day in a burrow. Night drives sometimes find one walking along or crossing the road.

SizeBody about 60–70 cm
DietRoots, fallen fruit, bark
ActiveNight
Where to lookWalking along the road
Part Two — The Wildlife23
Bornean Orangutan Pongo pygmaeus
Critically Endangered  ·  Diurnal  ·  regular

Deramakot's continued support of a healthy Orangutan population is one of the strongest arguments for the reduced-impact logging approach the reserve pioneered. A study modelling the reserve's management — "The Opportunity Cost of Engaging in Reduced-Impact Logging to Conserve the Orangutan" — found Deramakot held the highest Orangutan densities recorded among the Sabah forests compared, evidence that adequate regrowth periods between harvests and the deliberate retention of key food trees can outweigh the disturbance of controlled logging.

Sightings here are a daytime activity — a distant rustle of canopy resolving into a red-brown shape moving deliberately through the branches, or a fresh sleeping platform spotted from the road at dawn.

SizeBig males up to around 80 kg
DietFruit, young leaves, bark
ActiveDay
Where to lookFruiting trees; nests seen from the road
Part Two — The Wildlife24
Bornean Pygmy Elephant Elephas maximus borneensis
Endangered  ·  Diurnal / crepuscular  ·  occasional, unmistakable

The smallest and, by most accounts, gentlest subspecies of Asian elephant, found nowhere else in the world outside Borneo's northeast. Genetic studies suggest the population has been isolated long enough to be considered a distinct evolutionary lineage rather than a simple regional variant, with proportionally larger ears, a straighter tusk and a famously placid temperament compared to mainland relatives.

Deramakot sits within the wider Kinabatangan–Sabah landscape the population moves through, and herds move through the reserve rather than living there year-round. When they do, a sighting is hard to miss: browsed vegetation and fresh dung along the road often give notice before the animals themselves appear, usually as a loose family herd feeding unhurried at the treeline.

Part Two — The Wildlife25
Maroon Langur Presbytis rubicunda
Vulnerable  ·  Diurnal  ·  common

A striking, deep russet-red leaf monkey endemic to Borneo, typically encountered in small family troops moving through the mid-canopy on morning foraging rounds. Almost entirely folivorous, favouring young leaves and shoots.

Its habituation to vehicle traffic along Deramakot's main road makes it one of the more approachable primates in the reserve for daytime photography.

SizeSmall-dog size, long tail
DietYoung leaves, seeds
ActiveDay
Where to lookMid-canopy along the roads
Part Two — The Wildlife26
Southern Pig-tailed Macaque Macaca nemestrina
Endangered  ·  Diurnal  ·  common

Named for its short, upward-curled tail, this robust macaque moves through Deramakot in large, active troops, foraging on the ground and in the lower canopy for fruit, seeds and invertebrates.

Troops can number several dozen individuals, their movement often first announced by falling fruit husks and rustling understorey before any animal is visible.

SizeMedium-sized monkey, short tail
DietFruit, seeds, insects
ActiveDay
Where to lookOn the ground and in low trees near the road
Part Two — The Wildlife27
Part Two · The Wildlife
Two More Primates

Philippine Slow Loris

Nycticebus menagensis  ·  Vulnerable

A small, round-eyed primate in thick golden fur that climbs with slow, deliberate hand-over-hand grips. It feeds at night on insects, fruit and plant gum, and is one of the few venomous mammals: glands on its upper arms produce a toxin it mixes with saliva. Its bright orange eyeshine is one of the easier things to pick out on a night drive.

Bornean Gibbon

Hylobates funereus  ·  Endangered

Heard far more often than seen. Gibbon families sing loud, rising songs at dawn that carry for kilometres across the forest, one of the best sounds of an early morning at Deramakot. They travel through the canopy by swinging arm over arm, and each small family group holds its own territory.

Part Two — The Wildlife28
Part Two · The Wildlife
Squirrels of the Canopy

Sunda Giant Squirrel

Ratufa affinis

A big, bushy-tailed squirrel with cream underparts, a rusty head and a long banded tail. It is common at Deramakot and often seen by day, leaping from crown to crown with its tail streaming behind. The tail steadies the squirrel in the air and on thin branches.

Red Giant Flying Squirrel

Petaurista petaurista

A large nocturnal squirrel in rich chestnut fur, with a loose skin membrane between its front and back legs that turns it into a glider. It launches from the canopy and glides a long way to another trunk. Common on the night drives, and usually found by the glow of its eyes in the spotlight.

Part Two — The Wildlife29
Night falls at the forest headquarters, and the evening drive is about to begin.
Part Two · The Wildlife
Birds Around Camp

Red-bearded Bee-eater

Nyctyornis amictus

A heavy-set, forest bee-eater, green-bodied with a shaggy crimson throat. Unlike the open-country bee-eaters it sits quietly, sallying out for large flying insects. At Deramakot it is seen around the accommodation most days, so you rarely need to go looking for it.

Crested Serpent Eagle

Spilornis cheela

Deramakot's most conspicuous resident raptor, and there is nearly always one somewhere around the reception centre, perched on a branch or calling overhead with its far-carrying, ringing cry. A specialist hunter of snakes and other reptiles.

Part Two — The Wildlife31
Part Two · The Wildlife
Two More to Look For

Blyth's Paradise-flycatcher

Terpsiphone affinis

A striking flycatcher: adult males trail long white tail streamers and carry a glossy blue-black crest. It often poses well while you drive the roads, making it one of the easiest of the reserve's special birds to watch from the vehicle.

White-fronted Falconet

Microhierax latifrons

Found only in northern Borneo, and one of the smallest birds of prey in the world, black and white with a chestnut belly. It hunts large insects from an exposed perch, and this one is eyeing a weaver ant. Look for it on bare, open branches and wires, where its tiny size is easy to miss.

Part Two — The Wildlife32
Part Two · The Wildlife
Two Borneo Icons

Rhinoceros Hornbill

Buceros rhinoceros  ·  Vulnerable

Borneo's best-known bird, with a huge upturned orange-and-red casque on top of its bill. Pairs cross the canopy with loud, whooshing wingbeats, and their honking calls carry a long way. It depends on big old trees: the female seals herself into a tree hole to nest, and the male feeds her through a narrow slit.

Bornean Bristlehead

Pityriasis gymnocephala  ·  Near Threatened

Found nowhere but Borneo, and one of the birds visiting birders most want to see: a chunky black bird with a red head and a crown of short, bristly feathers. It moves through the canopy in small noisy groups, often with other species, and its calls are usually the first clue. Tall lowland forest like Deramakot's is the place to look.

Part Two — The Wildlife33
Part Two · The Wildlife
Frogs After Dark
Wallace's Flying Frog with large golden eyes and webbed feet, DeramakotJason Luke

Wallace's Flying Frog

Rhacophorus nigropalmatus

A large, bright green tree frog with dramatically enlarged, fully webbed feet that allow it to glide between trees. First documented by Alfred Russel Wallace during his own Bornean travels, it remains a favourite target for night walks, usually found on low vegetation near still water.

Harlequin Tree Frog

Rhacophorus pardalis

Another gliding frog, with big webbed feet and enormous eyes. Its colour ranges from greens to browns, with darker blotches along the flanks. Found at night on low branches and vegetation, usually not far from water.

Part Two — The Wildlife34
Part Two · The Wildlife
Snakes in the Spotlight

Green Vine Snake

Ahaetulla prasina

Thin as a pencil and almost invisible among leaves and vines, with a long pointed snout and a keyhole-shaped pupil. It sways like a tendril in the breeze and takes lizards and frogs. Mildly venomous, but not dangerous to people.

Dog-toothed Cat Snake hanging from a branch at night, DeramakotJason Luke

Dog-toothed Cat Snake

Boiga cynodon

A slender, night-active tree snake with big eyes and enlarged front teeth. By day it sleeps coiled in foliage; after dark it hunts birds, lizards and small mammals. Mildly venomous, and seen on night drives where the road runs under overhanging trees.

Reticulated Python

Malayopython reticulatus

One of the longest snakes in the world, patterned with an intricate net-like camouflage that gives the species its name. Found both on the forest floor and along the road edge, and occasionally swimming across the reserve's waterways.

Part Two — The Wildlife35
Part Three · Practical
Planning Your Visit

Getting There

Fly into Sandakan (SDK), then drive about two hours to the town of Telupid. Guests are met at Telupid at 2pm and taken into the reserve by 4WD, a further two to three hours. A private transfer from Sandakan can be arranged.

Accommodation

Guests stay at the forest headquarters in clean chalet-style rooms with air conditioning and hot water. A resident cook provides three meals a day, and dietary needs can be met with notice. Power is by generator, and wifi and phone signal are now available.

How Long to Stay

Two nights gives a first taste. Three to four nights gives a much better chance of the rarer species, and dedicated mammal watchers often stay a week.

Night Drives

Most visitors join an evening drive of two to four hours, spotlighting from an open vehicle. On specialist mammal-watching trips a night drive can run for seven to ten hours, through the small hours when several of the scarcer species are moving.

What to Bring

A head torch, binoculars, a light rain jacket, leech socks and a power bank for the occasional generator outage.

Night Drive Etiquette

  • Keep voices low and movements slow once an animal is spotted.
  • Leave the spotlight to your guide, and don't hold the beam on an animal longer than needed.
  • Multi-night stays substantially improve the odds for the reserve's rarer species.
Part Three — Practical36
Part Three · Practical
Field Checklist

620 species of lowland Sabah, grouped by family. Tick what you see, on the page or on a printed copy. Most are found across Sabah's lowland forests, and not every species here has been recorded in Deramakot itself.

Mammals178 species
Primates
Bornean OrangutanPongo pygmaeus
Proboscis MonkeyNasalis larvatus
Silvered LangurTrachypithecus cristatus
Red Leaf Monkey (Maroon Langur)Presbytis rubicunda
Long-tailed MacaqueMacaca fascicularis
Southern Pig-tailed MacaqueMacaca nemestrina
Bornean GibbonHylobates funereus
Western TarsierCephalopachus bancanus
Philippine Slow LorisNycticebus menagensis
Sabah Grey LangurPresbytis sabana
Carnivores
Sunda Clouded LeopardNeofelis diardi
Bornean Bay CatCatopuma badia
Leopard CatPrionailurus javanensis
Marbled CatPardofelis marmorata
Flat-headed CatPrionailurus planiceps
Sun BearHelarctos malayanus
Malay CivetViverra tangalunga
Island Palm CivetParadoxurus philippinensis
Masked Palm CivetPaguma larvata
Bornean Striped Palm CivetArctogalidia stigmatica
Banded Palm CivetHemigalus derbyanus
Otter CivetCynogale bennettii
BinturongArctictis binturong
Malay WeaselMustela nudipes
Yellow-throated MartenMartes flavigula
Hairy-nosed OtterLutra sumatrana
Smooth-coated OtterLutrogale perspicillata
Oriental Small-clawed OtterAonyx cinereus
Sunda Stink BadgerMydaus javanensis
Banded LinsangPrionodon linsang
Short-tailed MongooseHerpestes brachyurus
Collared MongooseHerpestes semitorquatus
Ungulates & Megafauna
Bornean ElephantElephas maximus borneensis
Bornean Bearded PigSus barbatus
Sambar DeerRusa unicolor
Red MuntjacMuntiacus muntjak
Bornean Yellow MuntjacMuntiacus atherodes
Greater Mouse-deerTragulus napu
Lesser Mouse-deerTragulus kanchil
BantengBos javanicus
Pangolin & Colugo
Sunda PangolinManis javanica
Sunda Colugo (Flying Lemur)Galeopterus variegatus
Squirrels
Sunda Giant SquirrelRatufa affinis
Prevost's SquirrelCallosciurus prevostii
Plantain SquirrelCallosciurus notatus
Black-banded SquirrelCallosciurus nigrovittatus
Low's SquirrelSundasciurus lowii
Horse-tailed SquirrelSundasciurus hippurus
Tufted Ground SquirrelRheithrosciurus macrotis
Red Giant Flying SquirrelPetaurista petaurista
Black Flying SquirrelAeromys tephromelas
Ear-spot SquirrelCallosciurus adamsi
Slender SquirrelSundasciurus tenuis
Jentink's SquirrelSundasciurus jentinki
Three-striped Ground SquirrelLariscus insignis
Four-striped Ground SquirrelLariscus hosei
Brooke's SquirrelSundasciurus brookei
Least Pygmy SquirrelExilisciurus exilis
Shrew-faced SquirrelRhinosciurus laticaudatus
Hose's Pygmy Flying SquirrelPetaurillus hosei
Lesser Pygmy Flying SquirrelPetaurillus emiliae
Thomas' Flying SquirrelAeromys thomasi
Horsefield's Flying SquirrelIomys horsfieldii
Grey-cheeked Flying SquirrelHylopetes lepidus
Hagen's Flying SquirrelPetinomys hageni
Whiskered Flying SquirrelPetinomys genibarbis
Vordermann's Flying SquirrelPetinomys vordermanni
Temminck's Flying SquirrelPetinomys setosus
Red-cheeked Flying SquirrelHylopetes spadiceus
Smoky Flying SquirrelPteromyscus pulverulentus
Spotted Giant Flying SquirrelPetaurista elegans
Treeshrews
Common TreeshrewTupaia glis
Lesser TreeshrewTupaia minor
Large TreeshrewTupaia tana
Pen-tailed TreeshrewPtilocercus lowii
Slender TreeshrewTupaia gracilis
Striped TreeshrewTupaia dorsalis
Shrews & Moonrats
MoonratEchinosorex gymnura
Asian House ShrewSuncus murinus
Pygmy White-toothed ShrewSuncus etruscus
Sunda ShrewCrocidura monticola
South Eastern White-toothed ShrewCrocidura fuliginosa
Part Three — Practical37
Field ChecklistSabah lowland wildlife
Rats, Mice & Porcupines
Long-tailed Giant RatLeopoldamys sabanus
Malaysian Field RatRattus tiomanicus
Malayan PorcupineHystrix brachyura
Thick-spined PorcupineHystrix crassispinis
Black RatRattus rattus
Ricefield RatRattus argentiventer
Polynesian RatRattus exulans
Brown RatRattus norvegicus
Muller's Giant Sunda RatSundamys muelleri
Dark-tailed Tree RatNiviventer cremoriventer
Rajah Spiny RatMaxomys rajah
Red Spiny RatMaxomys surifer
Chestnut-bellied Spiny RatMaxomys ochraceiventer
Small Spiny RatMaxomys baeodon
Whitehead's Spiny RatMaxomys whiteheadi
Grey Tree RatLenothrix canus
Asian House MouseMus castaneus
Indomalayan Pencil-tailed Tree MouseChiropodomys gliroides
Large Pencil-tailed Tree MouseChiropodomys major
Ranee MouseHaeromys margarettae
Long-tailed PorcupineTrichys fasciculata
Bats
Large Flying FoxPteropus vampyrus
Island Flying FoxPteropus hypomelanus
Cave Nectar BatEonycteris spelaea
Lesser Short-nosed Fruit BatCynopterus brachyotis
Diadem Roundleaf BatHipposideros diadema
Wrinkle-lipped Free-tailed BatChaerephon plicatus
Geoffroy's RousetteRousettus amplexicaudatus
Bare-backed RousetteRousettus spinalatus
Horsfield's Fruit BatCynopterus horsfieldii
Dusky Fruit BatPenthetor lucasi
Tailless Fruit BatMegaerops ecaudatus
Spot-winged Fruit BatBalionycteris maculata
Greater Nectar BatEonycteris major
Long-tongued Nectar BatMacroglossus minimus
Small Asian Sheath-tailed BatEmballonura alecto
Lesser Sheath-tailed BatEmballonura monticola
Black-bearded Tomb BatTaphozous melanopogon
Long-winged Tomb BatTaphozous longimanus
Lesser False Vampire BatMegaderma spasma
Malayan Slit-faced BatNycteris tragata
Bornean Horseshoe BatRhinolophus borneensis
Acuminate Horseshoe BatRhinolophus acuminatus
Creagh's Horseshoe BatRhinolophus creaghi
Large-eared Horseshoe BatRhinolophus philippinensis
Trefoil Horseshoe BatRhinolophus trifoliatus
Lesser Woolly Horseshoe BatRhinolophus sedulus
Dusky Leaf-nosed BatHipposideros ater
Bicoloured Roundleaf BatHipposideros bicolor
Ashy Roundleaf BatHipposideros cineraceus
Dayak Roundleaf BatHipposideros dyacorum
Borneo Roundleaf BatHipposideros doriae
Ridley's Leaf-nosed BatHipposideros ridleyi
Fawn Leaf-nosed BatHipposideros cervinus
Cantor's Roundleaf BatHipposideros galeritus
Wall Roosting Mouse-eared BatMyotis muricola
Peters's MyotisMyotis ater
Burmese Whiskered BatMyotis montivagus
Bats (continued)
Ridley's BatMyotis ridleyi
Horsfield's BatMyotis horsfieldii
Lesser Large-footed BatMyotis hasseltii
Java PipistrellePipistrellus javanicus
Least PipistrellePipistrellus tenuis
Kelaart's PipistrellePipistrellus ceylonicus
Red-brown PipistrelleHypsugo kitcheneri
Peters's PipistrelleFalsistrellus petersi
Brown PipistrelleHypsugo imbricatus
Big-eared PipistrelleHypsugo macrotis
Common Thick-thumbed BatGlischropus tylopus
Rohu's BatPhiletor brachypterus
False Serotine BatHesperoptenus doriae
Blanford's BatHesperoptenus blanfordi
Large False BatHesperoptenus tomesi
Greater Bamboo BatTylonycteris robustula
Lesser Bamboo BatTylonycteris pachypus
Lesser Asiatic Yellow BatScotophilus kuhlii
Round-eared Tube-nosed BatMurina cyclotis
Bronze Tube-nosed BatMurina aenea
Gilded Tube-nosed BatMurina rozendaali
Brown Tube-nosed BatMurina suilla
Papillose Woolly BatKerivoula papillosa
Hardwicke's Woolly BatKerivoula hardwickii
Clear-winged Woolly BatKerivoula pellucida
Small Woolly BatKerivoula intermedia
Least Woolly BatKerivoula minuta
Whitehead's Woolly BatKerivoula whiteheadi
Lenis Woolly BatKerivoula lenis
Peters's Trumpet-eared BatPhoniscus jagorii
Groove-toothed BatPhoniscus atrox
Western Bent-winged BatMiniopterus magnater
Common Bent-winged BatMiniopterus schreibersii
Small Bent-winged BatMiniopterus pusillus
Intermediate Long-fingered BatMiniopterus medius
Little Bent-winged BatMiniopterus australis
Naked BatCheiromeles torquatus
Malayan Free-tailed BatMops mops
Birds196 species
Pheasants & Partridges
Great ArgusArgusianus argus
Crested FirebackLophura ignita
Crestless FirebackLophura erythrophthalma
Red JunglefowlGallus gallus
Crested PartridgeRollulus rouloul
Chestnut-necklaced PartridgeArborophila charltonii
Bornean Peacock-PheasantPolyplectron schleiermacheri
Bulwer's PheasantLophura bulweri
Herons, Storks & Bitterns
Storm's StorkCiconia stormi
Lesser AdjutantLeptoptilos javanicus
Purple HeronArdea purpurea
Great-billed HeronArdea sumatrana
Striated HeronButorides striata
Yellow BitternIxobrychus sinensis
Black BitternIxobrychus flavicollis
Part Three — Practical38
Field ChecklistSabah lowland wildlife
Raptors
Wallace's Hawk-EagleNisaetus nanus
Blyth's Hawk-EagleNisaetus alboniger
Crested Serpent EagleSpilornis cheela
White-bellied Sea EagleHaliaeetus leucogaster
Lesser Fish EagleIchthyophaga humilis
Grey-headed Fish EagleIchthyophaga ichthyaetus
Rufous-bellied EagleLophotriorchis kienerii
Bat HawkMacheiramphus alcinus
Jerdon's BazaAviceda jerdoni
Black BazaAviceda leuphotes
Crested GoshawkAccipiter trivirgatus
BesraAccipiter virgatus
White-fronted FalconetMicrohierax latifrons
Changeable Hawk-EagleNisaetus cirrhatus
Black EagleIctinaetus malayensis
Rails
Slaty-breasted RailLewinia striata
White-breasted WaterhenAmaurornis phoenicurus
Pigeons & Doves
Green Imperial PigeonDucula aenea
Jambu Fruit DoveRamphiculus jambu
Little Green PigeonTreron olax
Cinnamon-headed Green PigeonTreron fulvicollis
Thick-billed Green PigeonTreron curvirostra
Pink-necked Green PigeonTreron vernans
Emerald DoveChalcophaps indica
Zebra DoveGeopelia striata
Spotted DoveSpilopelia chinensis
Large Green PigeonTreron capellei
Parrots & Parakeets
Blue-crowned Hanging-ParrotLoriculus galgulus
Blue-rumped ParrotPsittinus cyanurus
Long-tailed ParakeetPsittacula longicauda
Cuckoos & Malkohas
Chestnut-breasted MalkohaPhaenicophaeus curvirostris
Raffles's MalkohaRhinortha chlorophaea
Red-billed MalkohaZanclostomus javanicus
Black-bellied MalkohaPhaenicophaeus diardi
Chestnut-bellied MalkohaPhaenicophaeus sumatranus
Green-billed MalkohaPhaenicophaeus tristis
Greater CoucalCentropus sinensis
Lesser CoucalCentropus bengalensis
Violet CuckooChrysococcyx xanthorhynchus
Banded Bay CuckooCacomantis sonneratii
Square-tailed Drongo-CuckooSurniculus lugubris
Asian Emerald CuckooChrysococcyx maculatus
Bornean Ground-CuckooCarpococcyx radiatus
Owls, Nightjars & Frogmouths
Buffy Fish OwlKetupa ketupu
Barred Eagle-OwlBubo sumatranus
Brown Wood OwlStrix leptogrammica
Oriental Bay OwlPhodilus badius
Sunda Scops OwlOtus lempiji
Reddish Scops OwlOtus rufescens
Large FrogmouthBatrachostomus auritus
Owls, Nightjars & Frogmouths (continued)
Dulit FrogmouthBatrachostomus harterti
Gould's FrogmouthBatrachostomus stellatus
Sunda FrogmouthBatrachostomus cornutus
Large-tailed NightjarCaprimulgus macrurus
Malaysian Eared NightjarEurostopodus temminckii
Blyth's FrogmouthBatrachostomus affinis
Bonaparte's NightjarCaprimulgus concretus
Swifts & Treeswifts
Glossy SwiftletCollocalia esculenta
Edible-nest SwiftletAerodramus fuciphagus
Black-nest SwiftletAerodramus maximus
Silver-rumped NeedletailRhaphidura leucopygialis
Brown-backed NeedletailHirundapus giganteus
Whiskered TreeswiftHemiprocne comata
Grey-rumped TreeswiftHemiprocne longipennis
Mossy-nest SwiftletAerodramus salangana
Trogons
Diard's TrogonHarpactes diardii
Red-naped TrogonHarpactes kasumba
Scarlet-rumped TrogonHarpactes duvaucelii
Cinnamon-rumped TrogonHarpactes orrhophaeus
Orange-breasted TrogonHarpactes oreskios
Kingfishers, Bee-eaters & Rollers
Oriental Dwarf KingfisherCeyx erithacus
Rufous-collared KingfisherActenoides concretus
Banded KingfisherLacedo pulchella
Stork-billed KingfisherPelargopsis capensis
Ruddy KingfisherHalcyon coromanda
White-throated KingfisherHalcyon smyrnensis
Collared KingfisherTodiramphus chloris
Blue-eared KingfisherAlcedo meninting
Blue-banded KingfisherAlcedo euryzona
Blue-throated Bee-eaterMerops viridis
Red-bearded Bee-eaterNyctyornis amictus
DollarbirdEurystomus orientalis
Hornbills
Rhinoceros HornbillBuceros rhinoceros
Helmeted HornbillRhinoplax vigil
Bushy-crested HornbillAnorrhinus galeritus
Wrinkled HornbillRhabdotorrhinus corrugatus
White-crowned HornbillBerenicornis comatus
Black HornbillAnthracoceros malayanus
Oriental Pied HornbillAnthracoceros albirostris
Wreathed HornbillRhyticeros undulatus
Barbets & Woodpeckers
Red-crowned BarbetPsilopogon rafflesii
Golden-naped BarbetPsilopogon pulcherrimus
Yellow-crowned BarbetPsilopogon henricii
Gold-whiskered BarbetPsilopogon chrysopsis
Red-throated BarbetPsilopogon mystacophanos
Blue-eared BarbetPsilopogon duvaucelii
Brown BarbetCaloramphus fuliginosus
Great Slaty WoodpeckerMulleripicus pulverulentus
Buff-necked WoodpeckerMeiglyptes tukki
Buff-rumped WoodpeckerMeiglyptes tristis
Part Three — Practical39
Field ChecklistSabah lowland wildlife
Barbets & Woodpeckers (continued)
Orange-backed WoodpeckerChrysocolaptes validus
Maroon WoodpeckerBlythipicus rubiginosus
Rufous WoodpeckerMicropternus brachyurus
Checker-throated WoodpeckerChrysophlegma mentale
Banded WoodpeckerChrysophlegma miniaceum
Crimson-winged WoodpeckerPicus puniceus
White-bellied WoodpeckerDryocopus javensis
Rufous PiculetSasia abnormis
Malaysian HoneyguideIndicator archipelagicus
Broadbills & Pittas
Black-and-red BroadbillCymbirhynchus macrorhynchos
Banded BroadbillEurylaimus javanicus
Black-and-yellow BroadbillEurylaimus ochromalus
Green BroadbillCalyptomena viridis
Dusky BroadbillCorydon sumatranus
Bornean Banded PittaHydrornis schwaneri
Blue-headed PittaHydrornis baudii
Giant PittaHydrornis caeruleus
Black-crowned PittaErythropitta ussheri
Blue-banded PittaErythropitta arquata
Babblers
Chestnut-winged BabblerCyanoderma erythropterum
Black-throated BabblerStachyris nigricollis
Moustached BabblerMalacopteron magnirostre
Sooty-capped BabblerMalacopteron affine
Ferruginous BabblerMalacopteron bowdleri
Fluffy-backed Tit-BabblerMacronus ptilosus
Black-capped BabblerPellorneum capistratum
Short-tailed BabblerPellorneum malaccense
Horsfield's BabblerMalacocincla sepiaria
White-chested BabblerPellorneum rostratum
Chestnut-backed Scimitar BabblerPomatorhinus montanus
Rufous-crowned BabblerMalacopteron magnum
Grey-throated BabblerStachyris nigriceps
Black-throated Wren-BabblerGypsophila nigricollis
Bulbuls
Black-headed BulbulBrachypodius atriceps
Black-and-white BulbulBrachypodius melanoleucos
Buff-vented BulbulIole olivacea
Grey-cheeked BulbulAlophoixus bres
Yellow-vented BulbulPycnonotus goiavier
Olive-winged BulbulPycnonotus plumosus
Red-eyed BulbulPycnonotus brunneus
Spectacled BulbulPycnonotus erythropthalmos
Hairy-backed BulbulTricholestes criniger
Puff-backed BulbulPycnonotus eutilotus
Scaly-breasted BulbulIxos malaccensis
Cream-vented BulbulPycnonotus simplex
Ochraceous BulbulAlophoixus ochraceus
Flycatchers, Fantails & Shamas
Malaysian Blue FlycatcherCyornis turcosus
White-rumped ShamaCopsychus malabaricus
Rufous-tailed ShamaTrichixos pyrropygus
Rufous-chested FlycatcherFicedula dumetoria
Pied FantailRhipidura javanica
Spotted FantailRhipidura perlata
Drongos, Orioles & Cuckooshrikes
Greater Racket-tailed DrongoDicrurus paradiseus
Bronzed DrongoDicrurus aeneus
Crow-billed DrongoDicrurus annectens
Black-naped OrioleOriolus chinensis
Dark-throated OrioleOriolus xanthonotus
Bar-bellied CuckooshrikeCoracina striata
Lesser CuckooshrikeCoracina fimbriata
Scarlet MinivetPericrocotus speciosus
Pied TrillerLalage nigra
Sunbirds, Spiderhunters & Flowerpeckers
Ruby-cheeked SunbirdChalcoparia singalensis
Plain SunbirdAnthreptes simplex
Red-throated SunbirdAnthreptes rhodolaema
Purple-naped SunbirdHypogramma hypogrammicum
Crimson SunbirdAethopyga siparaja
Spectacled SpiderhunterArachnothera flavigaster
Long-billed SpiderhunterArachnothera robusta
Grey-breasted SpiderhunterArachnothera modesta
Orange-bellied FlowerpeckerDicaeum trigonostigma
Yellow-breasted FlowerpeckerPrionochilus maculatus
Scarlet-breasted FlowerpeckerPrionochilus thoracicus
Bornean FlowerpeckerDicaeum monticolum
Leafbirds, Ioras & Fairy-bluebird
Greater Green LeafbirdChloropsis sonnerati
Lesser Green LeafbirdChloropsis cyanopogon
Blue-winged LeafbirdChloropsis cochinchinensis
Common IoraAegithina tiphia
Green IoraAegithina viridissima
Asian Fairy-bluebirdIrena puella
Bornean Endemics & Miscellaneous
Bornean BristleheadPityriasis gymnocephala
Velvet-fronted NuthatchSitta frontalis
Reptiles174 species
Crocodiles
Saltwater CrocodileCrocodylus porosus
Malaysian False GharialTomistoma schlegelii
Turtles & Tortoises
Painted TerrapinBatagur borneoensis
Malaysian Box TurtleCuora amboinensis
Common Leaf TurtleCyclemys dentata
Sunda Leaf TurtleCyclemys enigmatica
Malaysian Giant TurtleOrlitia borneensis
Black Marsh TurtleSiebenrockiella crassicollis
Asian Giant TortoiseManouria emys
Asian Softshell TurtleAmyda cartilaginea
Malaysian Softshell TurtleDogania subplana
Asian Giant Softshell TurtlePelochelys cantorii
Agamid Lizards (Dragons)
Brown Shrub LizardAphaniotis fusca
Crested Green LizardBronchocela cristatella
Maned Forest LizardBronchocela jubata
Part Three — Practical40
Field ChecklistSabah lowland wildlife
Agamid Lizards (Dragons) (continued)
Garden LizardCalotes versicolor
Horned Flying LizardDraco cornutus
Crested Flying LizardDraco cristatellus
Fringed Flying LizardDraco fimbriatus
Red-bearded Flying LizardDraco haematopogon
Large Flying LizardDraco maximus
Black-bearded Flying LizardDraco melanopogon
Obscure Flying LizardDraco obscurus
Common Flying LizardDraco sumatranus
Bornean Angle-headed LizardGonocephalus borneensis
Doria's Angle-headed LizardGonocephalus doriae
Giant Angle-headed LizardGonocephalus grandis
Blue-eyed Angle-headed LizardGonocephalus liogaster
Bornean Shrub LizardPhoxophrys borneensis
Large-headed Shrub LizardPhoxophrys cephalus
Black-lipped Shrub LizardPhoxophrys nigrilabris
Geckos
Cat GeckoAeluroscalabotes felinus
Kinabalu Bent-toed GeckoCyrtodactylus baluensis
Peters's Bent-toed GeckoCyrtodactylus consobrinus
Inger's Bent-toed GeckoCyrtodactylus ingeri
Malayan Bent-toed GeckoCyrtodactylus malayanus
Yoshi's Bent-toed GeckoCyrtodactylus yoshii
Common Four-clawed GeckoGehyra mutilata
Tokay GeckoGekko gecko
Warty House GeckoGekko monarchus
Smith's Giant GeckoGekko smithii
Brooke's House GeckoHemidactylus brookii
Frilly Forest GeckoHemidactylus craspedotus
Asian House GeckoHemidactylus frenatus
Garnot's House GeckoHemidactylus garnotii
Frilly House GeckoHemidactylus platyurus
Common Worm GeckoHemiphyllodactylus typus
Brown's Camouflage GeckoLuperosaurus browni
Horsfield's Parachute GeckoPtychozoon horsfieldii
Kuhl's Parachute GeckoPtychozoon kuhli
Skinks & Other Lizards
Striped Bornean Tree SkinkApterygodon vittatum
Grey Tree SkinkDasia grisea
Olive Tree SkinkDasia olivacea
Mangrove SkinkEmoia atrocostata
Philippine Ground SkinkEutropis indeprensa
Common Sun SkinkEutropis multifasciata
Black-banded Ground SkinkEutropis rudis
Red-throated Ground SkinkEutropis rugifera
Nieuwenhuis's Tree SkinkLamprolepis nieuwenhuisii
Vyner's Tree SkinkLamprolepis vyneri
Bornean Striped SkinkLipinia inexpectata
Common Striped SkinkLipinia vittigera
Bampfylde's Giant SkinkLygosoma bampfyldei
Bowring's Supple SkinkLygosoma bowringii
Blue-throated Forest SkinkSphenomorphus cyanolaemus
Hallier's Forest SkinkSphenomorphus hallieri
Many-scaled Forest SkinkSphenomorphus multisquamatus
Sabah Forest SkinkSphenomorphus sabanus
Beccari's Water SkinkTropidophorus beccarii
Brooke's Water SkinkTropidophorus brookei
Small-footed Water SkinkTropidophorus micropus
Skinks & Other Lizards (continued)
Bornean Glass LizardOphisaurus buettikoferi
White-tailed Worm LizardDibamus leucurus
Inger's Worm LizardDibamus ingeri
Voris's Worm LizardDibamus vorisi
Long-tailed Grass LizardTakydromus sexlineatus
Monitor Lizards
Dumeril's Monitor LizardVaranus dumerilii
Rough-necked Monitor LizardVaranus rudicollis
Water MonitorVaranus salvator
Pythons, Blind & Primitive Snakes
Reticulated PythonMalayopython reticulatus
Bornean Short PythonPython breitensteini
Wart SnakeAcrochordus granulatus
Elephant Trunk SnakeAcrochordus javanicus
Common Pipe SnakeCylindrophis ruffus
Sunbeam SnakeXenopeltis unicolor
Leonard's Pipe SnakeAnomochilus leonardi
Brahminy Blind SnakeIndotyphlops braminus
Lined Blind SnakeRamphotyphlops lineatus
Muller's Blind SnakeTyphlops muelleri
Stoliczka's Stream SnakeStoliczkia borneensis
Colubrid Snakes (Rear-fanged & Non-venomous)
Speckle-headed Vine SnakeAhaetulla fasciolata
Green Vine SnakeAhaetulla prasina
Dog-toothed Cat SnakeBoiga cynodon
Mangrove Cat SnakeBoiga dendrophila
White-spotted Cat SnakeBoiga drapiezii
Jasper Cat SnakeBoiga jaspidea
Black-headed Cat SnakeBoiga nigriceps
Bicoloured Reed SnakeCalamaria battersbyi
Bornean Reed SnakeCalamaria borneensis
Grabowsky's Reed SnakeCalamaria grabowskyi
Hillenius's Reed SnakeCalamaria hilleniusi
Collared Reed SnakeCalamaria leucogaster
Low's Reed SnakeCalamaria lovii
Variable Reed SnakeCalamaria lumbricoidea
Prakke's Reed SnakeCalamaria prakkei
Red-headed Reed SnakeCalamaria schlegeli
Yellow-bellied Reed SnakeCalamaria suluensis
Short-tailed Reed SnakeCalamaria virgulata
Yellow-striped Trinket SnakeCoelognathus flavolineatus
Copper-headed Trinket SnakeCoelognathus radiatus
Beautiful Bronzeback Tree SnakeDendrelaphis formosus
Kopstein's Bronzeback Tree SnakeDendrelaphis kopsteini
Painted Bronzeback Tree SnakeDendrelaphis pictus
Striated Bronzeback Tree SnakeDendrelaphis striatus
Half-banded Bridled SnakeDryocalamus subannulatus
Three-banded Bridled SnakeDryocalamus tristigatus
Keel-bellied Whip SnakeDryophiops rubescens
Dark-grey Ground SnakeElapoidis fuscus
Orange-bellied SnakeGongylosoma baliodeirum
Striped Ground SnakeGongylosoma longicauda
Royal Tree SnakeGonyophis margaritatus
Red-tailed RacerGonyosoma oxycephalum
Tricoloured RingneckLiopeltis tricolor
Dusky Wolf SnakeLycodon albofuscus
Island Wolf SnakeLycodon capucinus
Part Three — Practical41
Field ChecklistSabah lowland wildlife
Colubrid Snakes (Rear-fanged & Non-venomous) (continued)
Brown Wolf SnakeLycodon effraenis
White-banded Wolf SnakeLycodon subcinctus
Jewelled Kukri SnakeOligodon everetti
Meyerink's Kukri SnakeOligodon meyerinkii
Eight-lined Kukri SnakeOligodon octolineatus
Mountain Reed SnakeOreocalamus hanitschi
White-collared Reed SnakePseudorabdion albonuchalis
Mocquard's Reed SnakePseudorabdion collaris
Dwarf Reed SnakePseudorabdion longiceps
Malayan Keeled Rat SnakePtyas carinata
Javan Rat SnakePtyas korros
White-bellied Rat SnakePtyas fusca
Striped Black-headed SnakeSibynophis geminatus
White-lipped Black-headed SnakeSibynophis melanocephalus
Triangled Black-headed SnakeSibynophis triangularis
Bornean Black SnakeStegonotus borneensis
Orange Brown SnakeXenelaphis ellipsifer
Malayan Brown SnakeXenelaphis hexagonotus
Pit Vipers
Sumatran Pit ViperParias sumatranus
Bornean Palm Pit ViperTrimeresurus borneensis
Bornean Keeled Green Pit ViperTropidolaemus subannulatus
Kraits, Coral Snakes & Cobras
Banded KraitBungarus fasciatus
Blue-headed KraitBungarus flaviceps
Blue Coral SnakeCalliophis bivirgatus
Malayan Striped Coral SnakeCalliophis intestinalis
Sumatran CobraNaja sumatrana
King CobraOphiophagus hannah
Water & Mud Snakes
Yellow-banded Mangrove SnakeCantoria violacea
Dog-faced Water SnakeCerberus rynchops
Rainbow Water SnakeEnhydris enhydris
Water & Mud Snakes (continued)
Grey Water SnakeEnhydris plumbea
Crab-eating Mangrove SnakeFordonia leucobalia
Puff-faced Water SnakeHomalopsis buccata
Keelbacks & Mock Vipers
White-fronted KeelbackAmphiesma flavifrons
Sarawak KeelbackAmphiesma sarawacense
Yellow-spotted Water SnakeHydrablabes periops
Blue-necked KeelbackMacropisthodon rhodomelas
Corrugated Water SnakeOpisthotropis typica
Painted Mock ViperPsammodynastes pictus
Mock ViperPsammodynastes pulverulentus
Speckle-bellied KeelbackRhabdophis chrysargos
Red-bellied KeelbackRhabdophis conspicillatus
Red-necked KeelbackRhabdophis subminiatus
Red-sided Keelback Water SnakeXenochrophis trianguligerus
Slug-eating Snakes
Blunt-headed Slug-eating SnakeAplopeltura boa
Smooth Slug-eating SnakeAsthenodipsas laevis
Malayan Slug-eating SnakeAsthenodipsas malaccanus
Keeled Slug-eating SnakePareas carinatus
Barred Slug-eating SnakePareas nuchalis
Amphibians72 species
Litter Frogs
Web-footed Dwarf Litter FrogLeptobrachella palmata
Rough-sided Dwarf Litter FrogLeptobrachella parva
Lowland Litter FrogLeptobrachium abbotti
Dring's Slender Litter FrogLeptolalax dringi
Painted Slender Litter FrogLeptolalax pictus
Horned Frogs
Rough Horned FrogMegophrys edwardinae
Bornean Horned FrogMegophrys nasuta
True Toads
Brown Slender ToadAnsonia leptopus
Long-fingered Slender ToadAnsonia longidigita
Spiny Slender ToadAnsonia spinulifer
River ToadPhrynoidis aspera
Giant River ToadPhrynoidis juxtasper
Crested ToadIngerophrynus divergens
Swamp ToadIngerophrynus quadriporcatus
Cross ToadLeptophryne borbonica
Brown Tree ToadPedostibes hosii
Marbled Tree ToadPedostibes rugosus
Lowland Dwarf ToadPelophryne signata
Narrow-mouthed Frogs
Blotch-sided Spadefoot FrogCalluella smithi
Saffron-bellied FrogChaperina fusca
Variable Sticky FrogKalophrynus heterochirus
Intermediate Sticky FrogKalophrynus intermedius
Rufous-sided Sticky FrogKalophrynus pleurostigma
Lesser Sticky FrogKalophrynus subterrestris
Brown BullfrogKaloula baleata
Tree Hole FrogMetaphrynella sundana
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Field ChecklistSabah lowland wildlife
Narrow-mouthed Frogs (continued)
Berdmore's Narrow-mouthed FrogMicrohyla berdmorei
Bornean Narrow-mouthed FrogMicrohyla borneensis
Pothole Narrow-mouthed FrogMicrohyla petrigena
True & Fanged Frogs
Mangrove FrogFejervarya cancrivora
Asian Grass FrogFejervarya limnocharis
Taiwanese FrogHoplobatrachus rugulosus
Hole-in-the-head FrogHuia cavitympanum
Inger's Dwarf FrogIngerana baluensis
Rough Guardian FrogLimnonectes finchi
Greater Swamp FrogLimnonectes ingeri
Kuhl's Creek FrogLimnonectes kuhlii
Giant River FrogLimnonectes leporinus
Peat Swamp FrogLimnonectes malesianus
Smooth Guardian FrogLimnonectes palavanensis
Lesser Swamp FrogLimnonectes paramacrodon
Seep FrogOccidozyga baluensis
Yellow-bellied Puddle FrogOccidozyga laevis
Stream & Rock Frogs
Northern Torrent FrogMeristogenys orphnocnemis
Brown Marsh FrogHylarana baramica
White-lipped FrogHylarana chalconota
Common Green FrogHylarana erythraea
Rough-sided FrogHylarana glandulosa
Malaysian FrogHylarana luctuosa
Cricket FrogHylarana nicobariensis
Spotted Stream FrogHylarana picturata
Striped Stream FrogHylarana signata
Hose's FrogOdorrana hosii
Rock Skipper FrogStaurois latopalmatus
Black-spotted Rock FrogStaurois natator
Green Spotted Rock FrogStaurois tuberilinguis
Tree & Flying Frogs
Cinnamon FrogNyctixalus pictus
Hose's Bush FrogPhilautus hosii
Collett's Tree FrogPolypedates colletti
Four-lined Tree FrogPolypedates leucomystax
Dark-eared Tree FrogPolypedates macrotis
File-eared Tree FrogPolypedates otilophus
Frilled Tree FrogRhacophorus appendiculatus
Blue-spotted Tree FrogRhacophorus cyanopunctatus
Jade Tree FrogRhacophorus dulitensis
Short-nosed Tree FrogRhacophorus gauni
Brown Tree FrogRhacophorus harrissoni
White-eared Tree FrogRhacophorus kajau
Wallace's Flying FrogRhacophorus nigropalmatus
Harlequin Tree FrogRhacophorus pardalis
Reinwardt's Flying FrogRhacophorus reinwardti
Red-legged Tree FrogRhacophorus rufipes
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Part Three · Practical
References & Acknowledgements
Photography and field observations of Deramakot's wildlife.
Jason Luke
Photographs of Wallace's Flying Frog and the Dog-toothed Cat Snake.
Tom Cooper
Aerial photography of Deramakot's base camp, forest canopy and roads.
Sabah Forestry Department
RIL and forest management documentation for Deramakot. forest.sabah.gov.my
Mammalwatching.com
Jon Hall's May 2015 trip report and later visitor accounts of the reserve's mammal community.
Lagan, P. et al., Ecological Research (2007)
"Sustainable use of tropical forests by reduced-impact logging in Deramakot Forest Reserve" — the reserve's RIL and FSC certification history.
Kate Humble (2019)
Her published account of a January 2019 visit to Deramakot, referenced in Part One.
Leibniz-IZW, Panthera & Sabah Forestry Department, Biotropica (2026)
Camera-trap study (2007–2023) on Sunda Clouded Leopard longevity and detection bias, referenced in the species profile.
Economics, Ecology & the Environment, WP 194
Orangutan density findings at Deramakot under reduced-impact logging, referenced in the species profile.
A book about Deramakot Forest Reserve, Sabah, Malaysian Borneo. Species information reflects natural-history knowledge at the time of writing. Conservation statuses follow the IUCN Red List and may since have changed.
Part Three — Practical44
The Mengaris — one of Borneo's tallest trees, often rising far above the surrounding canopy.
Deramakot

A working timber forest in the heart of Sabah, and one of the world's great places to watch wild mammals.

Plan your visit
deramakot.com
deramakotbookings@gmail.com
Night safaris and wildlife tours from 2 to 7 nights.
Adventure Alternative Borneo Sdn Bhd · Licensed tour operator KPL/LN 8442